Alexander,
The set of 4 additional antennas on the AP-335 is used to provide "antenna polarization diversity" for the 5GHz radio. On that radio, we basically connected two antennas to each radio chain (so 2 sets of four altogether). One antenna is vertically polarized (which is typical/default), the other is horizontally polarized. We use an RF switch to allow us to dynamically select the best antenna for each transmit and receive packet (software controlled).
There is no impact on MIMO support, but this allows us to potentially gain the equivalent of up to a couple of dB by optimizing the coupling between the antennas on the AP and client device.
Note that the gain and coverage patterns of the vertical and horizontal antenna elements are (roughly) the same.
The tricky part obviously is determing which set of four antenna elements to use (VVVV to HHHH) for each packet to get the best performance. We continue to improve and tweak the algorithm for that.
When using external antennas, this dynamic behavior is obviously lost, and both radios use a fixed set of 4 antenna elements each (or rather a total of 4 dual-band elements). There is no reduction in the 4x4:4 MIMO capabilities.
While this is an interesting and performance enhancing feature of the 335, the true value of the 330 Series access points (over the 320 and 310 series for example) is in the high platform performance and Smart Rate interface.